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Onthophagus {genus} -
Arthropoda
;
Insecta
;
Coleoptera
;
Scarabaeidae
;
Scarabaeinae
;
Onthophagini
;
Print
Search Taxonomy
Sub-taxa
Species (381)
Onthophagus AAX0250 [3]
Onthophagus AAX0301 [1]
Onthophagus ACS3363 [2]
Onthophagus ACS3364 [2]
Onthophagus ACS3774 [3]
Onthophagus ACS4435 [1]
Onthophagus ACS5398 [1]
Onthophagus acuminatus [46]
Onthophagus acuminatusAS1 [2]
Onthophagus acuminatusASolis01 [14]
Onthophagus acuminatusASolis02 [27]
Onthophagus acuminatusASolis03 [7]
Onthophagus aeruginosus [61]
Onthophagus aethiopicus [498]
Onthophagus aff. aeneus [1]
Onthophagus aff. andersoni [18]
Onthophagus aff. anthracinus [15]
Onthophagus aff. buculus [1]
Onthophagus aff. clypeatus [8]
Onthophagus aff. hircus [2]
Onthophagus aff. longimanus [12]
Onthophagus aff. nabeleki [1]
Onthophagus affin crinitus panamensis [2]
Onthophagus affin nabeleki [1]
Onthophagus alluvius [3]
Onthophagus alternans [1]
Onthophagus andalusicus [6]
Onthophagus andersoni [36]
Onthophagus andrewesi [1]
Onthophagus angulicollis [163]
Onthophagus anthracinus [22]
Onthophagus apterus [1]
Onthophagus arcifer [1]
Onthophagus asperulus [2]
Onthophagus atevimus [3]
Onthophagus atriglabrus [7]
Onthophagus atripennis [6]
Onthophagus atrofasciatus [39]
Onthophagus atrosericeus [6]
Onthophagus atrox [1]
Onthophagus auritus [1]
Onthophagus australis [7]
Onthophagus babirussa [5]
Onthophagus babirussoides [1]
Onthophagus baolocensis [1]
Onthophagus baraudi [2]
Onthophagus basipostulatus [2]
Onthophagus batesi [7]
Onthophagus bellus [3]
Onthophagus bengalensis [5]
Onthophagus bicallosus [7]
Onthophagus bicarinaticeps [1]
Onthophagus bicavicollis [2]
Onthophagus bidens [12]
Onthophagus bidentatus [10]
Onthophagus bifasciatus [1]
Onthophagus bindaree [2]
Onthophagus binodis [10]
Onthophagus birugatus [1]
Onthophagus bisectus [1]
Onthophagus bonasus [3]
Onthophagus boops [1]
Onthophagus borneensis [3]
Onthophagus breviconus [29]
Onthophagus brevifrons [4]
Onthophagus bronzeus [1]
Onthophagus browni [1]
Onthophagus buculus [2]
Onthophagus capella [5]
Onthophagus castetsi [1]
Onthophagus catharinensis [1]
Onthophagus catta [9]
Onthophagus centricornis [3]
Onthophagus cervicapra [1]
Onthophagus cervus [1]
Onthophagus cf. deflexicollis [1]
Onthophagus cf. jeannelianus [1]
Onthophagus cf. taurinus [1]
Onthophagus cf. tragus [1]
Onthophagus championi [16]
Onthophagus chryses [5]
Onthophagus clypeatus [3]
Onthophagus clypeatus group [2]
Onthophagus coccineus [2]
Onthophagus cochisus [4]
Onthophagus coenobita [50]
Onthophagus comatulus [1]
Onthophagus concinnus [5]
Onthophagus confusus [4]
Onthophagus consentaneus [1]
Onthophagus coriaceoumbrosus [6]
Onthophagus coscineus [10]
Onthophagus crassicollis [3]
Onthophagus cribripennis [1]
Onthophagus crinitus [12]
Onthophagus cruciger [2]
Onthophagus cryptodicranius [5]
Onthophagus cuniculus [1]
Onthophagus curvicornis [33]
Onthophagus cyanellus [129]
Onthophagus dama [1]
Onthophagus damaki [1]
Onthophagus dandalu [3]
Onthophagus dellacasai [2]
Onthophagus depressus [2]
Onthophagus diabolicus [1]
Onthophagus dicranius [8]
Onthophagus dicranoides [3]
Onthophagus digitifer [1]
Onthophagus dispar [28]
Onthophagus dorsipilulus [5]
Onthophagus dunningi [1]
Onthophagus duporti [1]
Onthophagus egrigius [3]
Onthophagus elegans [2]
Onthophagus emarginatus [2]
Onthophagus ensifer [9]
Onthophagus evanidus [1]
Onthophagus extensicollis [6]
Onthophagus falculatus [1]
Onthophagus fasciatus [2]
Onthophagus favrei [2]
Onthophagus ferox [1]
Onthophagus filicornis [16]
Onthophagus fimetarius [7]
Onthophagus fissicornis [2]
Onthophagus flavimargo [3]
Onthophagus fodiens [9]
Onthophagus fracticornis [43]
Onthophagus furcatus [11]
Onthophagus fuscopunctatus [1]
Onthophagus gazellinus [23]
Onthophagus genuinus [4]
Onthophagus gibbulus [12]
Onthophagus giraffa [1]
Onthophagus glabratus [1]
Onthophagus gracilipes [1]
Onthophagus granulatus [5]
Onthophagus grataehelenae [5]
Onthophagus griseosetosus [1]
Onthophagus grossepunctatus [5]
Onthophagus haagi [1]
Onthophagus haematopus [4]
Onthophagus hecate [42]
Onthophagus hinnulus [2]
Onthophagus hircus [1]
Onthophagus hirtus [4]
Onthophagus hopfneri [15]
Onthophagus humboldti [4]
Onthophagus illyricus [21]
Onthophagus incensus [21]
Onthophagus incensusASolis01 [4]
Onthophagus incensusASolis02 [49]
Onthophagus inediapterus [5]
Onthophagus insularis [1]
Onthophagus interstitialis [3]
Onthophagus joannae [21]
Onthophagus jugicola [24]
Onthophagus kondaoensis [2]
Onthophagus laevigatus [1]
Onthophagus laevissimus [11]
Onthophagus laminatus [2]
Onthophagus laminifer [1]
Onthophagus landolti [8]
Onthophagus landoltiASolis01 [2]
Onthophagus landoltiASolis02 [2]
Onthophagus landoltiASolis03 [1]
Onthophagus laticollis [1]
Onthophagus latigena [4]
Onthophagus lebasi [7]
Onthophagus lemur [11]
Onthophagus lenzi [1]
Onthophagus leroyi [82]
Onthophagus leucostigma [2]
Onthophagus limonensis [9]
Onthophagus lojanus [3]
Onthophagus longimanus [1]
Onthophagus lucidus [2]
Onthophagus madoqua [1]
Onthophagus maki [4]
Onthophagus malabarensis [1]
Onthophagus maniti [2]
Onthophagus marginalis [5]
Onthophagus marginicollis [11]
Onthophagus marginicollisAS1 [4]
Onthophagus marginicollisAS2 [9]
Onthophagus masaicus [3]
Onthophagus medius [83]
Onthophagus medorensis [3]
Onthophagus melitaeus [3]
Onthophagus merdarius [3]
Onthophagus micropterus [1]
Onthophagus millitaris [3]
Onthophagus minutus [1]
Onthophagus mirabilis [2]
Onthophagus mjobergi [1]
Onthophagus mulgravei [1]
Onthophagus nebulosus [1]
Onthophagus negligens [1]
Onthophagus nemorivagus [12]
Onthophagus nigriventris [3]
Onthophagus nitidior [2]
Onthophagus nr. babirussa [2]
Onthophagus nr. peninsularis [2]
Onthophagus nuchicornis [49]
Onthophagus nutans [1]
Onthophagus nyctopus [19]
Onthophagus obscurior [2]
Onthophagus ochromerus [1]
Onthophagus oklahomensis [8]
Onthophagus omostigma [84]
Onthophagus onorei [16]
Onthophagus onthochromus [5]
Onthophagus opacicollis [5]
Onthophagus orientalis [3]
Onthophagus orpheus [21]
Onthophagus orphnoides [6]
Onthophagus osculatii [14]
Onthophagus ovatus [45]
Onthophagus pactolus [4]
Onthophagus panici [2]
Onthophagus parvulus [1]
Onthophagus pedator [1]
Onthophagus penicillatus [6]
Onthophagus peninsulocupreus [3]
Onthophagus pennsylvanicus [10]
Onthophagus pentacanthus [1]
Onthophagus petenensis [6]
Onthophagus phanaeides [1]
Onthophagus polyodon [342]
Onthophagus polyphemi [1]
Onthophagus potanini [1]
Onthophagus praecellens [48]
Onthophagus propraecellens [16]
Onthophagus pseudovinctus [2]
Onthophagus pugionatus [29]
Onthophagus pullus [24]
Onthophagus punctatus [3]
Onthophagus pygmaeus [1]
Onthophagus quadridentatus [1]
Onthophagus quadrimaculata [101]
Onthophagus quetzalis [5]
Onthophagus rangifer [1]
Onthophagus rectecornutus [3]
Onthophagus rhinolophus [13]
Onthophagus rinophyllus [8]
Onthophagus rorarius [2]
Onthophagus rubicundulus [1]
Onthophagus rubrescens [14]
Onthophagus rubricatus [13]
Onthophagus rudis [3]
Onthophagus ruficapillus [12]
Onthophagus rugicollis [4]
Onthophagus rugulipennis [25]
Onthophagus sagittarius [8]
Onthophagus sansibaricus [1]
Onthophagus semiareus [1]
Onthophagus semiasper [16]
Onthophagus semicornis [11]
Onthophagus semifex [3]
Onthophagus seniculus [1]
Onthophagus sharpi [1]
Onthophagus sharpiASolis01 [1]
Onthophagus sharpiASolis02 [3]
Onthophagus sharpiASolis2 [1]
Onthophagus signatus [4]
Onthophagus similis [53]
Onthophagus simplex [119]
Onthophagus sinuosicollis [5]
Onthophagus sloanei [1]
Onthophagus socialis [1]
Onthophagus solisi [2]
Onthophagus solisiASolis01 [1]
Onthophagus solisiASolis02 [2]
Onthophagus solivagus [5]
Onthophagus sp [1]
Onthophagus sp. [14]
Onthophagus sp. 1 [1]
Onthophagus sp. 1 CWC-2007 [1]
Onthophagus sp. 1 SB-2018 [1]
Onthophagus sp. 2 [1]
Onthophagus sp. 2 CWC-2007 [1]
Onthophagus sp. 2 SB-2018 [1]
Onthophagus sp. 3 [1]
Onthophagus sp. 3 SB-2018 [1]
Onthophagus sp. 4 CWC-2007 [1]
Onthophagus sp. 4 SB-2018 [1]
Onthophagus sp. 5 CWC-2007 [1]
Onthophagus sp. 5 SB-2018 [1]
Onthophagus sp. 6 CWC-2007 [1]
Onthophagus sp. 6 SB-2018 [1]
Onthophagus sp. 7 SB-2018 [1]
Onthophagus sp.AAX0263 [2]
Onthophagus sp.ADV4499 [4]
Onthophagus sp.AFX5186 [4]
Onthophagus sp.AFX5219 [8]
Onthophagus sp.AFX6621 [7]
Onthophagus sp.AFX6932 [36]
Onthophagus sp.AFX6980 [1]
Onthophagus sp.AFX6982 [3]
Onthophagus sp.AFX6983 [2]
Onthophagus sp.AFX6986 [1]
Onthophagus sp.AFX6997 [1]
Onthophagus sp.AFX7002 [2]
Onthophagus sp.AFX7005 [1]
Onthophagus sp.AFX7007 [1]
Onthophagus sp.AFX7013 [1]
Onthophagus sp.AFX7014 [1]
Onthophagus sp.AFX7016 [1]
Onthophagus sp.AFX7026 [2]
Onthophagus sp.AGA8670 [8]
Onthophagus sp.AGA8672 [6]
Onthophagus sp.AGA8674 [2]
Onthophagus sp.AGA8675 [3]
Onthophagus sp. AMGS 80889 [1]
Onthophagus sp. AMGS 80890 [1]
Onthophagus sp. AMGS 80891 [1]
Onthophagus sp. AMGS 80896 [1]
Onthophagus sp. AMGS 80897 [1]
Onthophagus sp. AMGS 80900 [1]
Onthophagus sp. AMGS 80926 [1]
Onthophagus sp. AMGS 80931 [1]
Onthophagus sp. AMGS 80932 [1]
Onthophagus sp. BM670919 [1]
Onthophagus sp. BMNH669344 [1]
Onthophagus sp. BMNH670919 [1]
Onthophagus sp. BMNH676985 [1]
Onthophagus sp. BMNH676988 [1]
Onthophagus sp. BMNH676993 [1]
Onthophagus sp. BMNH677008 [1]
Onthophagus sp. NLG-2014 [1]
Onthophagus sp. SAEVG Morph0201 [1]
Onthophagus sp. SM-2012 [1]
Onthophagus sp. dgi1 [1]
Onthophagus sp. nov [36]
Onthophagus sp22 [1]
Onthophagus spinifex [1]
Onthophagus stigmosus [28]
Onthophagus stockwelli [29]
Onthophagus striatulus [11]
Onthophagus stylocerus [3]
Onthophagus subaeneus [1]
Onthophagus subtropicus [2]
Onthophagus sugillatus [1]
Onthophagus suturellus [2]
Onthophagus tapirus [7]
Onthophagus taurinus [3]
Onthophagus taurus [35]
Onthophagus teitanicus [40]
Onthophagus timetrius [1]
Onthophagus tonsus [17]
Onthophagus transisthmius [2]
Onthophagus transthsismius [9]
Onthophagus trapezicornis [10]
Onthophagus trigibber [2]
Onthophagus tristis [3]
Onthophagus tuberculifrons [8]
Onthophagus turbatus [1]
Onthophagus turneri [2]
Onthophagus unifasciatus [1]
Onthophagus usurpator [1]
Onthophagus vacca [89]
Onthophagus variegatus [1058]
Onthophagus variolaris [1]
Onthophagus velutinus [1]
Onthophagus vermiculatus [1]
Onthophagus verrucosus [11]
Onthophagus verticallis [7]
Onthophagus verticicornis [27]
Onthophagus viduus [3]
Onthophagus vinctus [1]
Onthophagus viridivinosus [5]
Onthophagus vitulus [10]
Onthophagus vividus [3]
Onthophagus vulpes [3]
Onthophagus xanthernerus [1]
Onthophagus xanthomerus [11]
Onthophagus xanthopterus [19]
Onthophagus yangi [1]
Onthophagus yucatanus [4]
Onthophagus yukae [1]
Taxon Description (Wikipedia)
full article at Wikipedia
BOLD Stats
Specimen Records:
5,482
Public Records:
1,547
Specimens with Sequences:
4,234
Public Species:
303
Specimens with Barcodes:
3,832
Public BINs:
252
Species:
381
Species With Barcodes:
289
Species List - Progress
Access Published & Released Data
Contributors (Specimens & Sequencing)
Specimen Depositories:
Sequencing Labs:
Imagery
images representing subtaxa of Onthophagus
(
Onthophagus yucatanus
)
Sample ID:
CRY070
License:
CreativeCommons Attribution Share-Alike (by-sa) (2016)
License Holder:
Colección de Insectos, El Colegio de la Frontera Sur, Unidad Tapachula
Collection Sites
Collected from 31 countries.
Top 20:
Show All Countries
Costa Rica
561
Colombia
75
Malaysia
20
Honduras
171
Pakistan
63
United Kingdom
15
Germany
121
Canada
59
South Africa
14
Australia
115
Kenya
55
Netherlands
12
Austria
89
Peru
51
Argentina
10
United States
79
Mexico
27
India
7
Ecuador
77
France
26
Taxon Occurrence (GBIF)
![(Onthophagus AAX0250 - QCAZI_122077) @13 [ ] Unspecified (default): All Rights Reserved (2014) David A. Donoso & Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/122077_Dorsal%2B1412023440.JPG "(Onthophagus AAX0250 - QCAZI_122077) {} [ ] Unspecified (default): All Rights Reserved (2014)")
![(Onthophagus AAX0301 - QCAZI_121973) @13 [ ] CreativeCommons - Attribution (2014) David A. Donoso & Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121973_Dorsal%2B1411416582.JPG "(Onthophagus AAX0301 - QCAZI_121973) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus ACS3363 - QCAZI_121989) @13 [ ] CreativeCommons - Attribution (2014) David A. Donoso & Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121989_Dorsal%2B1411413300.JPG "(Onthophagus ACS3363 - QCAZI_121989) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus ACS3364 - QCAZI_121991) @14 [ ] CreativeCommons - Attribution (2014) David A. Donoso and Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121991_Dorsal%2B1411413736.JPG "(Onthophagus ACS3364 - QCAZI_121991) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus ACS3774 - QCAZI_121924) @14 [ ] CreativeCommons - Attribution (2014) David A. Donoso & Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121924_Dorsal%2B1412781738.JPG "(Onthophagus ACS3774 - QCAZI_121924) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus ACS4435 - QCAZI_121977) @14 [ ] CreativeCommons - Attribution (2014) David A. Donoso and Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121977_Dorsal%2B1411414376.JPG "(Onthophagus ACS4435 - QCAZI_121977) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus ACS5398 - QCAZI_121978) @13 [ ] CreativeCommons - Attribution (2014) David A. Donoso and Emilia Moreno PUCE](http://131.104.63.36/pics/_w70/FFECU/121978_Ventral%2B1411416338.JPG "(Onthophagus ACS5398 - QCAZI_121978) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus acuminatus - KMP_DB_50) @11 [ ] CreativeCommons Attribution Non-Commercial (by-nc (2024) Unspecified Technische Universität Darmstadt](http://131.104.63.36/pics/_w70/ASFL/Ont_acu_DB50_D_01%2B1721925564.jpg "(Onthophagus acuminatus - KMP_DB_50) {} [ ] CreativeCommons Attribution Non-Commercial (by-nc (2024)")
![(Onthophagus acuminatusAS1 - IAvH-E-151437) @13 [ ] CreativeCommons - Attribution Non-Commercial (2015) Instituto Humboldt IAvH Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt](http://131.104.63.36/pics/_w70/IHCOL/IMG_2215%2B1437600176.JPG "(Onthophagus acuminatusAS1 - IAvH-E-151437) {} [ ] CreativeCommons - Attribution Non-Commercial (2015)")
![(Onthophagus acuminatusASolis01 - INB0004102890) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004102890%2B1310141754.jpg "(Onthophagus acuminatusASolis01 - INB0004102890) {} [ ] Copyright (2010)")
![(Onthophagus acuminatusASolis02 - INB0004186077) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004186077%2B1310141942.jpg "(Onthophagus acuminatusASolis02 - INB0004186077) {} [ ] Copyright (2010)")
![(Onthophagus acuminatusASolis03 - ASolis001829) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2022) Bernardo Espinoza Guanacaste Dry Forest Conservation Fund](http://131.104.63.36/pics/_w70/CRBAF/ASolis001829%2B1670527848.jpg "(Onthophagus acuminatusASolis03 - ASolis001829) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2022)")
![(Onthophagus aeruginosus - BIOUG46475-G05) @14 [ ] CreativeCommons - Attribution (2020) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/KMPUF/BIOUG46475-G05%2B1597451922.jpg "(Onthophagus aeruginosus - BIOUG46475-G05) {} [ ] CreativeCommons - Attribution (2020)")
![(Onthophagus aethiopicus - MPALA-OPP-HYENA-02) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/MPALA-OPP-HYENA-02_V1%2B1706891239.jpg "(Onthophagus aethiopicus - MPALA-OPP-HYENA-02) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus aff. aeneus - IBS019) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Instituto de Biología Subtropical Instituto de Biología Subtropical](http://131.104.63.36/pics/_w70/IBSCG/IBS019%2B1537326444.jpg "(Onthophagus aff. aeneus - IBS019) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus aff. andersoni - BIOUG28574-C08) @11 [ ] CreativeCommons - Attribution (2018) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNPDB/BIOUG28574-C08%2B1528141950.jpg "(Onthophagus aff. andersoni - BIOUG28574-C08) {} [ ] CreativeCommons - Attribution (2018)")
![(Onthophagus aff. anthracinus - BIOUG28575-C09) @11 [ ] CreativeCommons - Attribution (2018) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNPDB/BIOUG28575-C09%2B1528819298.jpg "(Onthophagus aff. anthracinus - BIOUG28575-C09) {} [ ] CreativeCommons - Attribution (2018)")
![(Onthophagus aff. buculus - IBS089) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Instituto de Biología Subtropical Instituto de Biología Subtropical](http://131.104.63.36/pics/_w70/IBSCG/IBS089%2B1537326268.jpg "(Onthophagus aff. buculus - IBS089) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus aff. clypeatus - RE21-KP1- COL-0021) @11 [ ] CreativeCommons Attribution Non-Commercial (by-nc (2024) Unspecified Technische Universität Darmstadt](http://131.104.63.36/pics/_w70/ASFL/Ont_cly_12_D_001%2B1721055578.jpg "(Onthophagus aff. clypeatus - RE21-KP1- COL-0021) {} [ ] CreativeCommons Attribution Non-Commercial (by-nc (2024)")
![(Onthophagus aff. hircus - IBS018) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Instituto de Biología Subtropical Instituto de Biología Subtropical](http://131.104.63.36/pics/_w70/IBSCG/IBS018%2B1537326446.jpg "(Onthophagus aff. hircus - IBS018) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus aff. longimanus - BIOUG28574-G07) @11 [ ] CreativeCommons - Attribution (2018) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNPDB/BIOUG28574-G07%2B1528213374.jpg "(Onthophagus aff. longimanus - BIOUG28574-G07) {} [ ] CreativeCommons - Attribution (2018)")
![(Onthophagus aff. nabeleki - MECN_EN_39532) @11 [ ] CC-BY-NC-ND (2024) INABIO INABIO](http://131.104.63.36/pics/_w70/BIOEC/MECN_EN_39532_dorsal%2B1733345122.JPG "(Onthophagus aff. nabeleki - MECN_EN_39532) {} [ ] CC-BY-NC-ND (2024)")
![(Onthophagus affin crinitus panamensis - MECN_EN_41159) @11 [ ] CC-BY-NC-ND (2024) INABIO INABIO](http://131.104.63.36/pics/_w70/BIOEC/MECN_EN_41159_dorsal%2B1733774996.JPG "(Onthophagus affin crinitus panamensis - MECN_EN_41159) {} [ ] CC-BY-NC-ND (2024)")
![(Onthophagus affin nabeleki - MECN_EN_39531) @11 [ ] CC-BY-NC-ND (2024) INABIO INABIO](http://131.104.63.36/pics/_w70/BIOEC/MECN_EN_39531_dorsal%2B1733344980.JPG "(Onthophagus affin nabeleki - MECN_EN_39531) {} [ ] CC-BY-NC-ND (2024)")
![(Onthophagus alluvius - CNC COLEO 00119933) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNCCB/CNC_COLEO_00119933%2B1308767472.JPG "(Onthophagus alluvius - CNC COLEO 00119933) {} [ ] CreativeCommons - Attribution (2011)")
![(Onthophagus alternans - CCDB-34064-A11) @11 [ ] CreativeCommons - Attribution (2019) CBG Photography Group Smithsonian Institution](http://131.104.63.36/pics/_w70/SICOD/CCDB-34064-A11%2B1565709558.jpg "(Onthophagus alternans - CCDB-34064-A11) {} [ ] CreativeCommons - Attribution (2019)")
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![(Onthophagus anthracinus - INB0003797921) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003797921%2B1310141658.jpg "(Onthophagus anthracinus - INB0003797921) {} [ ] Copyright (2010)")
![(Onthophagus arcifer - SPEC#4741) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%234741_V1%2B1706913002.jpg "(Onthophagus arcifer - SPEC#4741) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus atrofasciatus - SPEC#3881) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%233881_V1%2B1706891232.jpg "(Onthophagus atrofasciatus - SPEC#3881) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus cruciger - IB_330) @11 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/IBS/IB_330%2B1660329752.jpg "(Onthophagus cruciger - IB_330) {} [ ] Unspecified (default): All Rights Reserved")
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![(Onthophagus egrigius - UMKL-TG006) @13 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/FIBM/tg006%2B1340128626.jpg "(Onthophagus egrigius - UMKL-TG006) {} [ ] Unspecified (default): All Rights Reserved")
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![(Onthophagus gibber - SPEC#4217) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%234217_V1%2B1706891268.jpg "(Onthophagus gibber - SPEC#4217) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus granulatus - UNEIECOL00017) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike Unspecified Unspecified](http://131.104.63.36/pics/_w70/IECOL/UNECOL17D%2B1533148400.jpg "(Onthophagus granulatus - UNEIECOL00017) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike")
![(Onthophagus grataehelenae - INB0004179643) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004179643%2B1310141942.jpg "(Onthophagus grataehelenae - INB0004179643) {} [ ] Copyright (2010)")
![(Onthophagus grossepunctatus - BGE_00102_B02) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/PC210303%2B1706314398.JPG "(Onthophagus grossepunctatus - BGE_00102_B02) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023)")
![(Onthophagus hecate - 09BBCOL-0255) @15 [ ] CreativeCommons - Attribution (2009) Unspecified Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/USCOL/IMG_8335%2B1250520100.JPG "(Onthophagus hecate - 09BBCOL-0255) {} [ ] CreativeCommons - Attribution (2009)")
![(Onthophagus hirtus - BGE_00590_D09) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/P5131595%2B1723305438.JPG "(Onthophagus hirtus - BGE_00590_D09) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024)")
![(Onthophagus hopfneri - INB0003303354) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003303354%2B1310141564.jpg "(Onthophagus hopfneri - INB0003303354) {} [ ] Copyright (2010)")
![(Onthophagus humboldti - INB0004134630) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004134630%2B1310141846.jpg "(Onthophagus humboldti - INB0004134630) {} [ ] Copyright (2010)")
![(Onthophagus illyricus - BC ZSM COL 01421) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010) Stefan Schmidt SNSB, Zoologische Staatssammlung Muenchen](http://131.104.63.36/pics/_w70/FBCOB/BCZSMCOL01421%2B1283415252.JPG "(Onthophagus illyricus - BC ZSM COL 01421) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2010)")
![(Onthophagus incensus - INB0003356979) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003356979%2B1310141566.jpg "(Onthophagus incensus - INB0003356979) {} [ ] Copyright (2010)")
![(Onthophagus incensusASolis01 - ASolis000584) @11 [ ] CreativeCommons Attribution NonCommercial ShareAlike (2021) Angel Solis BioAlfa](http://131.104.63.36/pics/_w70/CRBAF/ASolis000584%2B1632939142.jpg "(Onthophagus incensusASolis01 - ASolis000584) {} [ ] CreativeCommons Attribution NonCommercial ShareAlike (2021)")
![(Onthophagus incensusASolis02 - INB0004206593) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004206593%2B1310141942.jpg "(Onthophagus incensusASolis02 - INB0004206593) {} [ ] Copyright (2010)")
![(Onthophagus inediapterus - INB0003112175) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003112175%2B1310141468.jpg "(Onthophagus inediapterus - INB0003112175) {} [ ] Copyright (2010)")
![(Onthophagus insularis - MUTPL-CISEC-0047794) @11 [ ] CYC (2024) UTPL UTPL](http://131.104.63.36/pics/_w70/BEELO/MUTPL-CISEC-0047794%2B1724427293.jpeg "(Onthophagus insularis - MUTPL-CISEC-0047794) {} [ ] CYC (2024)")
![(Onthophagus joannae - BFB_Col_FK_5623) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) Zoologische Staatssammlung Muenchen SNSB, Zoologische Staatssammlung Muenchen](http://131.104.63.36/pics/_w70/FBCOG/BC_ZSM_COL_05623%2B1338575756.JPG "(Onthophagus joannae - BFB_Col_FK_5623) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Onthophagus jugicola - CCDB-34064-E12) @11 [ ] CreativeCommons - Attribution (2019) CBG Photography Group Smithsonian Institution](http://131.104.63.36/pics/_w70/SICOD/CCDB-34064-E12%2B1565720002.jpg "(Onthophagus jugicola - CCDB-34064-E12) {} [ ] CreativeCommons - Attribution (2019)")
![(Onthophagus kondaoensis - BGE_00900_F06) @11 [ ] Copyright (c) (2025) Zinenko Oleksandr Museum of Nature of V.N. Karazin Kharkiv National University](http://131.104.63.36/pics/_w70/TSKNU/F6_Onthophagus_kondaoensis%2B315518400.JPG "(Onthophagus kondaoensis - BGE_00900_F06) {} [ ] Copyright (c) (2025)")
![(Onthophagus laevissimus - SPEC#6914) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236914_V1%2B1706912824.jpg "(Onthophagus laevissimus - SPEC#6914) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus laminatus - gvc3683-1L) @12 [ ] Copyright (2004) Unspecified Unspecified](http://131.104.63.36/pics/_w70/COQT/gvc3683a%2B1113258782.jpg "(Onthophagus laminatus - gvc3683-1L) {} [ ] Copyright (2004)")
![(Onthophagus laminifer - MUTPL-CISEC-0047976) @11 [ ] CYC (2024) UTPL UTPL](http://131.104.63.36/pics/_w70/BEELO/MUTPL-CISEC-0047976%2B1724378945.jpeg "(Onthophagus laminifer - MUTPL-CISEC-0047976) {} [ ] CYC (2024)")
![(Onthophagus landolti - IAvH-E-161320) @13 [ ] CreativeCommons - Attribution Non-Commercial (2015) Instituto Humboldt IAvH Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt](http://131.104.63.36/pics/_w70/IHCOL/IMG_2264%2B1437664692.JPG "(Onthophagus landolti - IAvH-E-161320) {} [ ] CreativeCommons - Attribution Non-Commercial (2015)")
![(Onthophagus landoltiASolis01 - INB0003077453) @14 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003077453%2B1310141468.jpg "(Onthophagus landoltiASolis01 - INB0003077453) {} [ ] Copyright (2010)")
![(Onthophagus landoltiASolis02 - INB0004207462) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004207462%2B1310141942.jpg "(Onthophagus landoltiASolis02 - INB0004207462) {} [ ] Copyright (2010)")
![(Onthophagus landoltiASolis03 - INB0003354637) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003354637%2B1310141564.jpg "(Onthophagus landoltiASolis03 - INB0003354637) {} [ ] Copyright (2010)")
![(Onthophagus latigena - BGE_00102_B04) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/PC210307%2B1706314404.JPG "(Onthophagus latigena - BGE_00102_B04) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023)")
![(Onthophagus lebasi - IAvH-E-161315) @11 [ ] CreativeCommons - Attribution Non-Commercial (2015) Instituto Humboldt IAvH Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt](http://131.104.63.36/pics/_w70/IHCOL/IMG_2258%2B1437663730.JPG "(Onthophagus lebasi - IAvH-E-161315) {} [ ] CreativeCommons - Attribution Non-Commercial (2015)")
![(Onthophagus lemur - GBOLRL0116) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://131.104.63.36/pics/_w70/GBMUS/GBOLRL0116%2B1502467510.JPG "(Onthophagus lemur - GBOLRL0116) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
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![(Onthophagus limonensis - INB0004186091) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004186091%2B1310141942.jpg "(Onthophagus limonensis - INB0004186091) {} [ ] Copyright (2010)")
![(Onthophagus lojanus - MUTPL-CISEC-0041852) @11 [ ] CYC (2024) UTPL UTPL](http://131.104.63.36/pics/_w70/BEELO/MUTPL-CISEC-0041852%2B1724426529.jpeg "(Onthophagus lojanus - MUTPL-CISEC-0041852) {} [ ] CYC (2024)")
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![(Onthophagus marginicollis - INB0004207583) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004207583%2B1310141942.jpg "(Onthophagus marginicollis - INB0004207583) {} [ ] Copyright (2010)")
![(Onthophagus marginicollisAS1 - IAvH-E-151444) @12 [ ] CreativeCommons - Attribution Non-Commercial (2015) Instituto Humboldt IAvH Instituto de Investigacion de Recursos Biologicos Alexander von Humboldt](http://131.104.63.36/pics/_w70/IHCOL/IMG_2216%2B1437600226.JPG "(Onthophagus marginicollisAS1 - IAvH-E-151444) {} [ ] CreativeCommons - Attribution Non-Commercial (2015)")
![(Onthophagus masaicus - CCDB-37899-C06) @11 [ ] CreativeCommons - Attribution Share - Alike (2024) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/KENCO/CCDB-37899-C06%2B1709225080.jpg "(Onthophagus masaicus - CCDB-37899-C06) {} [ ] CreativeCommons - Attribution Share - Alike (2024)")
![(Onthophagus medius - BFB_Col_FK_8745) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) Unspecified SNSB, Zoologische Staatssammlung Muenchen](http://131.104.63.36/pics/_w70/FBCOG/BFB_Col_FK_8745%2B1343938442.jpg "(Onthophagus medius - BFB_Col_FK_8745) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Onthophagus medorensis - CNC COLEO 00119950) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNCCB/CNC_COLEO_00119950%2B1308767840.JPG "(Onthophagus medorensis - CNC COLEO 00119950) {} [ ] CreativeCommons - Attribution (2011)")
![(Onthophagus melitaeus - BGE_00102_B09) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/PC210321%2B1706314420.JPG "(Onthophagus melitaeus - BGE_00102_B09) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2023) (2023)")
![(Onthophagus merdarius - BGE_00590_C04) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/P5131536%2B1723305386.JPG "(Onthophagus merdarius - BGE_00590_C04) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024)")
![(Onthophagus micropterus - INBIOCRI002378988) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI002378988%2B1310142130.jpg "(Onthophagus micropterus - INBIOCRI002378988) {} [ ] Copyright (2010)")
![(Onthophagus millitaris - SPEC#2925) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%232925_V1%2B1706617369.jpg "(Onthophagus millitaris - SPEC#2925) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus mirabilis - MECN_EN_22866) @11 [ ] CC-BY-NC-ND (2024) INABIO INABIO](http://131.104.63.36/pics/_w70/BIOEC/MECN_EN_22866_dorsal%2B1733421852.JPG "(Onthophagus mirabilis - MECN_EN_22866) {} [ ] CC-BY-NC-ND (2024)")
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![(Onthophagus nemorivagus - INB0003879164) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003879164%2B1310141658.jpg "(Onthophagus nemorivagus - INB0003879164) {} [ ] Copyright (2010)")
![(Onthophagus nuchicornis - BIOUG09885-C12) @15 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/INRMA/BIOUG09885-C12%2B1390854096.jpg "(Onthophagus nuchicornis - BIOUG09885-C12) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus nyctopus - INBIOCRI002410004) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI002410004%2B1310142130.jpg "(Onthophagus nyctopus - INBIOCRI002410004) {} [ ] Copyright (2010)")
![(Onthophagus oklahomensis - CCDB-23512-A05) @14 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/LYMAB/CCDB-23512-A05%2B1441289606.jpg "(Onthophagus oklahomensis - CCDB-23512-A05) {} [ ] CreativeCommons - Attribution (2015)")
![(Onthophagus omostigma - SPEC#6078) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236078_V1%2B1706891261.jpg "(Onthophagus omostigma - SPEC#6078) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus onorei - EBSQ_BC154) @11 [ ] by-nc-nd (2033) Unspecified Estación Biológica Manu](http://131.104.63.36/pics/_w70/PSCAB/IMG_6530%2B1677015872.JPG "(Onthophagus onorei - EBSQ_BC154) {} [ ] by-nc-nd (2033)")
![(Onthophagus onthochromus - EBLA_BC292) @11 [ ] by-nc-nd (2033) Unspecified Estación Biológica Manu](http://131.104.63.36/pics/_w70/PSCAB/EBLA_BC292%2B1677101032.JPG "(Onthophagus onthochromus - EBLA_BC292) {} [ ] by-nc-nd (2033)")
![(Onthophagus opacicollis - LPRC-JT21-072) @11 [ ] Creative common (2022) Julien Touroult Museum national d'Histoire naturelle, Paris](http://131.104.63.36/pics/_w70/LPRCI/LPRCJT21_072%2B1643194165.jpg "(Onthophagus opacicollis - LPRC-JT21-072) {} [ ] Creative common (2022)")
![(Onthophagus orpheus - CNC COLEO 00119964) @14 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNCCB/CNC_COLEO_00119964%2B1308768880.JPG "(Onthophagus orpheus - CNC COLEO 00119964) {} [ ] CreativeCommons - Attribution (2011)")
![(Onthophagus orphnoides - INBIOCRI002465999) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI002465999%2B1310142224.jpg "(Onthophagus orphnoides - INBIOCRI002465999) {} [ ] Copyright (2010)")
![(Onthophagus osculatii - BC-GF2024-0757) @11 [ ] Creative Commons (2024) rodolphe rougerie Museum national d'histoire naturelle](http://131.104.63.36/pics/_w70/FGBEL/GF_0757%2B315525600.JPG "(Onthophagus osculatii - BC-GF2024-0757) {} [ ] Creative Commons (2024)")
![(Onthophagus ovatus - GBOL_Col_FK_1983) @13 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015) Unspecified SNSB, Zoologische Staatssammlung Muenchen](http://131.104.63.36/pics/_w70/GBCOC/GBOL_Col_FK_1983%2B1343851694.jpg "(Onthophagus ovatus - GBOL_Col_FK_1983) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2015)")
![(Onthophagus pactolus - NIBGE COL-01212) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://131.104.63.36/pics/_w70/MACOL/NIBGE_COL-1212%2B1308121280.JPG "(Onthophagus pactolus - NIBGE COL-01212) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Onthophagus panici - BGE_00590_C09) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/P5131552%2B1723305401.JPG "(Onthophagus panici - BGE_00590_C09) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024)")
![(Onthophagus penicillatus - UMKL-TG002) @11 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/FIBM/tg002%2B1340128170.jpg "(Onthophagus penicillatus - UMKL-TG002) {} [ ] Unspecified (default): All Rights Reserved")
![(Onthophagus peninsulocupreus - UMKL-TG001) @12 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/FIBM/tg001%2B1340128020.jpg "(Onthophagus peninsulocupreus - UMKL-TG001) {} [ ] Unspecified (default): All Rights Reserved")
![(Onthophagus pennsylvanicus - CCDB-23512-A10) @13 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/LYMAB/CCDB-23512-A10%2B1441724528.JPG "(Onthophagus pennsylvanicus - CCDB-23512-A10) {} [ ] CreativeCommons - Attribution (2015)")
![(Onthophagus polyodon - SPEC#4300) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%234300_V1%2B1706891046.jpg "(Onthophagus polyodon - SPEC#4300) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus polyphemi - CCDB-23512-A12) @11 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/LYMAB/CCDB-23512-A12%2B1441724860.JPG "(Onthophagus polyphemi - CCDB-23512-A12) {} [ ] CreativeCommons - Attribution (2015)")
![(Onthophagus potanini - NIBGE COL-01179) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011) Muhammad Ashfaq National Institute for Biotechnology and Genetic Engineering Faisalabad Pakistan](http://131.104.63.36/pics/_w70/MACOL/NIBGE_COL-1179%2B1308107534.JPG "(Onthophagus potanini - NIBGE COL-01179) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2011)")
![(Onthophagus praecellens - INB0004207496) @14 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0004207496%2B1310141942.jpg "(Onthophagus praecellens - INB0004207496) {} [ ] Copyright (2010)")
![(Onthophagus propraecellens - INB0003981789) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003981789%2B1310141754.jpg "(Onthophagus propraecellens - INB0003981789) {} [ ] Copyright (2010)")
![(Onthophagus pseudovinctus - CCDB-34064-H10) @11 [ ] CreativeCommons - Attribution (2019) CBG Photography Group Smithsonian Institution](http://131.104.63.36/pics/_w70/SICOD/CCDB-34064-H10%2B1565724284.jpg "(Onthophagus pseudovinctus - CCDB-34064-H10) {} [ ] CreativeCommons - Attribution (2019)")
![(Onthophagus pugionatus - CCDB-34064-F05) @11 [ ] CreativeCommons - Attribution (2019) CBG Photography Group Smithsonian Institution](http://131.104.63.36/pics/_w70/SICOD/CCDB-34064-F05%2B1565720568.jpg "(Onthophagus pugionatus - CCDB-34064-F05) {} [ ] CreativeCommons - Attribution (2019)")
![(Onthophagus pullus - SPEC#1581) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%231581_V1%2B1706828644.jpg "(Onthophagus pullus - SPEC#1581) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus punctatus - BGE_00104_H01) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/P1110755%2B1706316744.JPG "(Onthophagus punctatus - BGE_00104_H01) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024)")
![(Onthophagus quadrimaculata - SPEC#6122) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236122_V1%2B1706890996.jpg "(Onthophagus quadrimaculata - SPEC#6122) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus quetzalis - INBIOCRI000768258) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI000768258%2B1310142036.jpg "(Onthophagus quetzalis - INBIOCRI000768258) {} [ ] Copyright (2010)")
![(Onthophagus raffrayi - SPEC#6118) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236118_V1%2B1706890966.jpg "(Onthophagus raffrayi - SPEC#6118) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus rhinolophus - BIOUG28572-F03) @11 [ ] CreativeCommons - Attribution (2018) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNPDB/BIOUG28572-F03%2B1529073290.JPG "(Onthophagus rhinolophus - BIOUG28572-F03) {} [ ] CreativeCommons - Attribution (2018)")
![(Onthophagus rinophyllus - EBLA_BC301) @11 [ ] by-nc-nd (2033) Unspecified Estación Biológica Manu](http://131.104.63.36/pics/_w70/PSCAB/EBLA_BC301%2B1677101032.JPG "(Onthophagus rinophyllus - EBLA_BC301) {} [ ] by-nc-nd (2033)")
![(Onthophagus rubrescens - EBLA_BC296) @11 [ ] by-nc-nd (2033) Unspecified Estación Biológica Manu](http://131.104.63.36/pics/_w70/PSCAB/EBLA_BC296%2B1677101032.JPG "(Onthophagus rubrescens - EBLA_BC296) {} [ ] by-nc-nd (2033)")
![(Onthophagus rubricatus - SPEC#8168) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%238168_V1%2B1707472860.jpg "(Onthophagus rubricatus - SPEC#8168) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus rudis - UMKL-TG013) @12 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/FIBM/tg013%2B1340129350.jpg "(Onthophagus rudis - UMKL-TG013) {} [ ] Unspecified (default): All Rights Reserved")
![(Onthophagus ruficapillus - ABOL-BioBlitz21-0509) @11 [ ] CC BY-SA (2021) Rabl, Dominik Unspecified](http://131.104.63.36/pics/_w70/TDAOE/21_0509%2B1647627174.jpg "(Onthophagus ruficapillus - ABOL-BioBlitz21-0509) {} [ ] CC BY-SA (2021)")
![(Onthophagus rugulipennis - SPEC#0618) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%230618_V1%2B1706828706.jpg "(Onthophagus rugulipennis - SPEC#0618) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus sagittarius - gvc14240-1L) @13 [ ] Copyright (2004) Unspecified Research Collection of Graeme Cocks](http://131.104.63.36/pics/_w70/COQT/gvc14240%2B1271717078.jpg "(Onthophagus sagittarius - gvc14240-1L) {} [ ] Copyright (2004)")
![(Onthophagus semiasper - SPEC#6357) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236357_V1%2B1706891437.jpg "(Onthophagus semiasper - SPEC#6357) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus semicornis - GBOLRL0117) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016) SNSB, Staatliche Naturwissenschaftliche Sammlungen Bayerns ZSM (SNSB, Zoologische Staatssammlung Muenchen)](http://131.104.63.36/pics/_w70/GBMUS/GBOLRL0117%2B1502467512.JPG "(Onthophagus semicornis - GBOLRL0117) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2016)")
![(Onthophagus semifex - UMKL-TG008) @11 [ ] Unspecified (default): All Rights Reserved Unspecified Unspecified](http://131.104.63.36/pics/_w70/FIBM/tg008%2B1340128978.jpg "(Onthophagus semifex - UMKL-TG008) {} [ ] Unspecified (default): All Rights Reserved")
![(Onthophagus sharpiASolis01 - INBIOCRI000641540) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI000641540%2B1310142036.jpg "(Onthophagus sharpiASolis01 - INBIOCRI000641540) {} [ ] Copyright (2010)")
![(Onthophagus sharpiASolis02 - INB0003127433) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003127433%2B1310141468.jpg "(Onthophagus sharpiASolis02 - INB0003127433) {} [ ] Copyright (2010)")
![(Onthophagus sharpiASolis2 - INBIOCRI000864372) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI000864372%2B1310142036.jpg "(Onthophagus sharpiASolis2 - INBIOCRI000864372) {} [ ] Copyright (2010)")
![(Onthophagus similis - RMNH.INS.542779) @14 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013) Unspecified Naturalis Biodiversity Center](http://131.104.63.36/pics/_w70/NLCOC/RMNH.INS.542779%2B1445576390.jpg "(Onthophagus similis - RMNH.INS.542779) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2013)")
![(Onthophagus simplex - SPEC#8034) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%238034_V1%2B1706891218.jpg "(Onthophagus simplex - SPEC#8034) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus sinuosicollis - SPEC#3024) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/3024_3x%2B1695126268.jpg "(Onthophagus sinuosicollis - SPEC#3024) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus solisi - INBIOCRI001287055) @11 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INBIOCRI001287055%2B1310142130.jpg "(Onthophagus solisi - INBIOCRI001287055) {} [ ] Copyright (2010)")
![(Onthophagus solisiASolis01 - INB0003879142) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003879142%2B1310141658.jpg "(Onthophagus solisiASolis01 - INB0003879142) {} [ ] Copyright (2010)")
![(Onthophagus solisiASolis02 - INB0003879276) @13 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003879276%2B1310141658.jpg "(Onthophagus solisiASolis02 - INB0003879276) {} [ ] Copyright (2010)")
![(Onthophagus sp - JBWM0314047) @11 [ ] Copyright (2008) Unspecified J.B. Wallis Museum of Entomology, University of Manitoba](http://131.104.63.36/pics/_w70/DSCOM/314047%2B978332400.JPG "(Onthophagus sp - JBWM0314047) {} [ ] Copyright (2008)")
![(Onthophagus sp. 1 - UAIC1021072) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Wendy Moore University of Arizona](http://131.104.63.36/pics/_w70/PIMA/UAIC1021072_Onthophagus_sp1_65mm2x_compedit%2B1530907026.jpg "(Onthophagus sp. 1 - UAIC1021072) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus sp. 2 - UAIC1023824) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Wendy Moore University of Arizona](http://131.104.63.36/pics/_w70/PIMA/UAIC1023824_Onthophagus_sp2_65mm2x_compedit%2B1530907107.jpg "(Onthophagus sp. 2 - UAIC1023824) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus sp. 3 - UAIC1022308) @11 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018) Wendy Moore University of Arizona](http://131.104.63.36/pics/_w70/PIMA/UAIC1022308_Onthophagus_sp3_65mm2x_compedit%2B1530907196.jpg "(Onthophagus sp. 3 - UAIC1022308) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2018)")
![(Onthophagus sp.AAX0263 - SPEC#6817) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236817_V1%2B1706890996.jpg "(Onthophagus sp.AAX0263 - SPEC#6817) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus sp.AFX5186 - MPALA-T2-ELE-05) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/Mpala_T2_ELE_05_18_Jul_2021_1680_mins_3x%2B1695615976.jpg "(Onthophagus sp.AFX5186 - MPALA-T2-ELE-05) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus sp.AFX7013 - SPEC#6248) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%236248_V1%2B1706891161.jpg "(Onthophagus sp.AFX7013 - SPEC#6248) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus sp.AFX7014 - SPEC#4728) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%234728_V1%2B1706891211.jpg "(Onthophagus sp.AFX7014 - SPEC#4728) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus striatulus - CNC COLEO 00119970) @11 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNCCB/CNC_COLEO_00119970%2B1308768782.JPG "(Onthophagus striatulus - CNC COLEO 00119970) {} [ ] CreativeCommons - Attribution (2011)")
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![(Onthophagus subaeneus - CCDB-23512-B06) @11 [ ] CreativeCommons - Attribution (2015) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/LYMAB/CCDB-23512-B06%2B1441725510.JPG "(Onthophagus subaeneus - CCDB-23512-B06) {} [ ] CreativeCommons - Attribution (2015)")
![(Onthophagus subtropicus - CNC COLEO 00119971) @13 [ ] CreativeCommons - Attribution (2011) CNC/CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/CNCCB/CNC_COLEO_00119971%2B1308768760.JPG "(Onthophagus subtropicus - CNC COLEO 00119971) {} [ ] CreativeCommons - Attribution (2011)")
![(Onthophagus suturellus - BGE_00590_C06) @11 [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024) Research Center in Biodiversity and Genetic Resources Research Center in Biodiversity and Genetic Resources](http://131.104.63.36/pics/_w70/BBIOP/P5131542%2B1723305392.JPG "(Onthophagus suturellus - BGE_00590_C06) {} [ ] Creative Commons Attribution NonCommercial ShareAlike (2024) (2024)")
![(Onthophagus tapirus - INB0003573260) @12 [ ] Copyright (2010) A. Solis Instituto Nacional de Biodiversidad](http://131.104.63.36/pics/_w70/ASSCR/INB0003573260%2B1310141566.jpg "(Onthophagus tapirus - INB0003573260) {} [ ] Copyright (2010)")
![(Onthophagus taurus - BIOUG09885-D03) @14 [ ] CreativeCommons - Attribution (2014) CBG Photography Group Centre for Biodiversity Genomics](http://131.104.63.36/pics/_w70/INRMA/BIOUG09885-D03%2B1390854332.JPG "(Onthophagus taurus - BIOUG09885-D03) {} [ ] CreativeCommons - Attribution (2014)")
![(Onthophagus teitanicus - SPEC#4843) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%234843_V1%2B1706890926.jpg "(Onthophagus teitanicus - SPEC#4843) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
![(Onthophagus timetrius - TEO-11-0103) @12 [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012) Mamadi Theresa Sethusa University of Johannesburg](http://131.104.63.36/pics/_w70/SAFIN/TEO-11-0103_1%2B1342447800.JPG "(Onthophagus timetrius - TEO-11-0103) {} [ ] CreativeCommons - Attribution Non-Commercial Share-Alike (2012)")
![(Onthophagus tonsus - SPEC#5917) @11 [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024) Unspecified Princeton University](http://131.104.63.36/pics/_w70/MRCDB/SPEC%235917_V1%2B1706891479.jpg "(Onthophagus tonsus - SPEC#5917) {} [ ] CreativeCommons Attribution Non-Commercial Share-Alike (2024)")
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![(Onthophagus trapezicornis - CCDB-34064-H08) @11 [ ] CreativeCommons - Attribution (2019) CBG Photography Group Smithsonian Institution](http://131.104.63.36/pics/_w70/SICOD/CCDB-34064-H08%2B1565724060.jpg "(Onthophagus trapezicornis - CCDB-34064-H08) {} [ ] CreativeCommons - Attribution (2019)")
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